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Soil moisture is a primary driver of comammox Nitrospira ab..:

Chisholm, C ; Di, Hong ; Cameron, Keith...
The original publication is available from Elsevier B.V. - https://doi.org/10.1016/j.scitotenv.2022.159961 - http://dx.doi.org/10.1016/j.scitotenv.2022.159961.  , 2023