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pnp-1 mutants have increased expression of IPR genes:

Eillen Tecle (10676052) ; Crystal B. Chhan (10676055) ; Latisha Franklin (10676058)...
https://figshare.com/articles/figure/_i_pnp-1_i_mutants_have_increased_expression_of_IPR_genes_/14455551.  , 2021