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From analysis of protein structural alignments toward a nov..:

Sunyaev, Shamil R. ; Bogopolsky, Gennady A. ; Oleynikova, Natalia V....
Proteins: Structure, Function, and Bioinformatics.  54 (2003)  3 - p. 569-582 , 2003
Link: https://doi.org/10.1002/prot.10503