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Proton uptake by cytochrome c oxidase on reduction and on l..:

Mitchell, Roy ; Rich, Peter R.
Biochimica et Biophysica Acta (BBA) - Bioenergetics.  1186 (1994)  1-2 - p. 19-26 , 1994
Link: https://doi.org/10.1016/0005-2728(94)90130-9