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Aging, microglia and cytoskeletal regulation are key factor..:

Janssen, Leen ; Dubbelaar, Marissa L. ; Holtman, Inge R....
Biochimica et Biophysica Acta (BBA) - Molecular Basis of Disease.  1863 (2017)  2 - p. 395-405 , 2017
Link: https://doi.org/10.1016/j.bbadis.2016.11.014