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Contribution of the residue at position 4 within classical ..:

Smith, Kate M. ; Di Antonio, Veronica ; Bellucci, Luca...
Biochimica et Biophysica Acta (BBA) - Molecular Cell Research.  1865 (2018)  8 - p. 1114-1129 , 2018
Link: https://doi.org/10.1016/j.bbamcr.2018.05.006