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Is His54 a gating residue for the ferritin ferroxidase site:

Bernacchioni, Caterina ; Ciambellotti, Silvia ; Theil, Elizabeth C..
Biochimica et Biophysica Acta (BBA) - Proteins and Proteomics.  1854 (2015)  9 - p. 1118-1122 , 2015
Link: https://doi.org/10.1016/j.bbapap.2015.02.011