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Cathepsin K inhibition renders equine bone marrow nucleated..:

Hussein, Hayam ; Boyaka, Prosper ; Dulin, Jennifer.
Comparative Immunology, Microbiology and Infectious Diseases.  45 (2016)  - p. 40-47 , 2016
Link: https://doi.org/10.1016/j.cimid.2016.02.005