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Ectoparasites are unlikely to be a primary cause of populat..:

Holz, Peter H. ; Lumsden, Linda F. ; Hufschmid, Jasmin
International Journal for Parasitology: Parasites and Wildlife.  7 (2018)  3 - p. 423-428 , 2018
Link: https://doi.org/10.1016/j.ijppaw.2018.10.006