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A symbiosome membrane is not required for the actions of tw..:

Grant, A.J. ; Trautman, D.A. ; Frankland, S..
Comparative Biochemistry and Physiology Part A: Molecular & Integrative Physiology.  135 (2003)  2 - p. 337-345 , 2003
Link: https://doi.org/10.1016/s1095-6433(03)00081-3