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Simultaneous amino acid substitutions at antigenic sites dr..:

Shih, Arthur Chun-Chieh ; Hsiao, Tzu-Chang ; Ho, Mei-Shang.
Proceedings of the National Academy of Sciences.  104 (2007)  15 - p. 6283-6288 , 2007
Link: https://doi.org/10.1073/pnas.0701396104