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Alpaca (Vicugna pacos), the first nonprimate species with a..:

Fichtner, Alina S. ; Karunakaran, Mohindar M. ; Gu, Siyi...
Proceedings of the National Academy of Sciences.  117 (2020)  12 - p. 6697-6707 , 2020
Link: https://doi.org/10.1073/pnas.1909474117