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Orientation of the LexA DNA-binding motif on operator DNA a..:

Dumoulin, P ; Oertel-Buchheit, P ; Granger-Schnarr, M.
Proceedings of the National Academy of Sciences.  90 (1993)  5 - p. 2030-2034 , 1993
Link: https://doi.org/10.1073/pnas.90.5.2030