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Both DNA gyrase and reverse gyrase are present in the hyper..:

Guipaud, Olivier ; Marguet, Evelyne ; Noll, Kenneth M...
Proceedings of the National Academy of Sciences.  94 (1997)  20 - p. 10606-10611 , 1997
Link: https://doi.org/10.1073/pnas.94.20.10606