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Heterosis in the second and third generation affects litter..:

Thirstrup, Janne ; Pertoldi, Cino ; Larsen Foged, Peter.
Archives of Biological Sciences; Arhiv za bioloske nauke.  66 (2014)  3 - p. 1097-1103 , 2014
Link: https://doi.org/10.2298/abs1403097t