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SUMO2 and SUMO3 redundantly prevent a noncanonical type I i..:

Crowl, John T. ; Stetson, Daniel B.
Proceedings of the National Academy of Sciences of the United States of America.  115 (2018)  26 - p. 6798-6803 , 2018
Link: https://www.jstor.org/stable/26510817