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Eyespots deflect predator attack increasing fitness and pro..:

Prudic, Kathleen L. ; Stoehr, Andrew M. ; Wasik, Bethany R..
Proceedings of the Royal Society B: Biological Sciences.  282 (2015)  1798 - p. 20141531 , 2015
Link: https://doi.org/10.1098/rspb.2014.1531